Graduate Education

Pathology and Anatomical Integrative Anatomy

Students interested in the study of Evolutionary and/or Integrative Anatomy (IA) may apply specifically to this focus area within the Pathobiology Area Program. Our graduate training emphasizes a flexible interdisciplinary approach to the education of doctoral students, with a current and growing core of closely-knit faculty in the Department of Pathology and Anatomical Sciences at the University of Missouri School of Medicine.

The University of Missouri (MU) is unique in affording access to a wide variety of resources on the same campus, some of which include the School of Medicine, College of Arts and Sciences, College of Engineering, College of Veterinary Medicine and Bond Life Sciences Center. This proximity facilitates access to exceptionally diverse and cutting-edge resources, with myriad cross-disciplinary opportunities for translational and evolutionary research and education.

In combination with the technical and theoretical proficiency of IA faculty, we capitalize on emerging approaches from allied programs in pathobiology and the life sciences. Campus-wide, there are numerous faculty and student groups with exciting research interests that complement those of IA. If you are interested in obtaining a PhD in Integrative Anatomy, please contact the Director of Graduate Studies before applying online.

Dr. Kevin Middleton
Associate Professor & Director of Graduate Studies
Department of Pathology & Anatomical Sciences
M263 Medical Sciences Building
University of Missouri School of Medicine
One Hospital Drive
Columbia, MO 65212

phone: 573-884-3192
fax: 573-884-4612

Degree Requirements

Integrative Anatomy PhD students are required to complete the minimum requirements of the MU Graduate School. These include 72 semester hours beyond the baccalaureate degree, including research and readings credits, to be approved by the doctoral advisor. Up to 30 hours of post-baccalaureate credit from an accredited institution may be transferred toward the doctoral degree. Further details are available online from the MU Graduate School. They also take courses in human gross anatomy, evolutionary biology, evolutionary morphology, statistics, and the ethical conduct of research, as well as participating in journal club, outreach activities, and teaching. Students must pass comprehensive exams and defend a research proposal, and must complete and defend a written dissertation.

Admission Requirements

  • Undergraduate and Post-Graduate Transcripts
  • Letters of recommendation: 3 required
  • Statement of Purpose/Intent
  • Curriculum vitae (or résumé)

Application Procedures

Applications for admissions are due yearly on the second Friday of January for Fall semester matriculation. Applications are not reviewed at other times.

Submit via the online application to the University of Missouri Graduate School:

  • All required Graduate School and PAS documents, such as transcripts, GRE scores (Pathology
     or Anatomy 
    ), TOEFL or IELTS scores (if required; code 6875). Although the MU Graduate School does not require GRE scores, these scores are required for programs within PAS.
  • Statement of purpose
  • Letters of recommendation
  • 3 letters of recommendation are required. Provide these names and contact information for each in the "Recommendation Provider List".
  • Curriculum vitae or resume
  • At least one writing sample (e.g., publications, proposals, upper-level course term papers, thesis, etc.)

Important: When registering online, choose the following options

  • For "Admissions Category": choose Doctorate- On Campus
  • For "Program of Interest": choose Pathobiology Area Program/Integrative Anatomy Focus/Doctor of Philosophy (PhD)